1. CIRAD, UMR RPB, 911 Avenue Agropolis, B.P. 64501, 34394 Montpellier Cedex 5, France; Centro Nacional de Investigaciones de Café (Cenicafé), A.A. 2427 Manizales, Caldas, Colombia; IRD, UMR DIAPC, and IRD, UMR RPB, 911 Avenue Agropolis, B.P. 64501, 34394 Montpellier Cedex 5, France