1. Graduate School of Pharmaceutical Sciences, The University of Tokyo, Bunkyo-ku, Tokyo 113-0033, Japan
2. School of Pharmaceutical Sciences, South-Central Minzu University, Wuhan 430074, China
3. Collaborative Research Institute for Innovative Microbiology, The University of Tokyo, Bunkyo-ku, Tokyo 113-0033, Japan
4. RIKEN Center for Sustainable Resource Science, Wako, Saitama 351-0198, Japan