1. State Key Laboratory of Microbial Technology, Shandong University, Qingdao, Shandong 266237, China
2. Shandong Provincial Key Laboratory of Synthetic Biology, CAS Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, No. 189 Songling Road, Qingdao, Shandong 266101, China
3. University of Chinese Academy of Sciences, No. 19(A) Yuquan Road, Beijing 100049, China
4. College of Chemistry and Pharmaceutical Sciences, Qingdao Agricultural University, Qingdao, Shandong 266109, China