1. Contribution from the Research Institute for Science and Engineering, Waseda University, 3-4-1 Okubo, Shinjuku-ku, Tokyo 169-8555, Japan, PRESTO, Japan Science and Technology Agency, 4-1-8 Honcho, Kawagichi-shi, Saitama 332-0012, Japan, and Division of Cell and Molecular Biology, Faculty of Natural Sciences, Imperial College London, South Kensington Campus, London SW7 2AZ, United Kingdom