1. Guangdong Key Laboratory of Fermentation and Enzyme Engineering, School of Biology and Biological Engineering, South China University of Technology, Guangzhou 510006, P. R. China
2. Guangdong research center of Industrial enzyme and Green manufacturing technology, School of Biology and Biological Engineering, South China University of Technology, Guangzhou 510006, P. R. China
3. Department of Biology, Massachusetts Institute of Technology, Cambridge, Massachusetts 02139, United States
4. Institute of Food Safety and Nutrition, Jinan University, Guangzhou 510632, P. R. China