1. Department of Botany and Microbiology, Faculty of Science, Beni Suef University, 62521 Beni-Suef, Egypt
2. Biology Department, Faculty of Science Yanbu, Taibah University, King Khalid Road, Al Amoedi, 46423 Yanbu El-Bahr, Saudi Arabia
3. Department of Botany and Microbiology, Faculty of Science, Cairo University, Giza 12613, Egypt
4. Laboratory for Molecular Plant Physiology and Biotechnology, Department of Biology, University of Antwerp, Groenenborgerlaan 171, B-2020, Antwerp, Belgium