1. National and Local United Engineering Research Center of Industrial Microbiology and Fermentation Technology; College of Life Sciences, Fujian Normal University, Fuzhou 350117, Fujian, People’s Republic of China
2. CAS Key Laboratory of Microbial Physiological and Metabolic Engineering, Institute of Microbiology, Chinese Academy of Sciences, No. 1 West Beichen Road, Chaoyang District, Beijing 100101, China