1. State Key Laboratory of Biological Fermentation Engineering of Beer, R&D Center, Tsingtao Brewery Company Ltd., Qingdao 266101, Shandong, PR China
2. State Key Laboratory of Food Nutrition and Safety, Tianjin University of Science & Technology, Tianjin 300457, PR China
3. Key Laboratory of Industrial Fermentation Microbiology (Ministry of Education), Tianjin University of Science & Technology, Tianjin 300457, PR China