1. Key Laboratory of Tibetan Medicine Research, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, 810008, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. State Key Laboratory of Plateau Ecology and Agriculture, Qinghai University, Xining 810016, China
4. Qinghai Provincial Key Laboratory of Tibetan Medicine Research, Xining, 810001, People’s Republic of China