1. Botanical Institute, Christian-Albrechts University Kiel, Olshausenstr. 40, Kiel D-24098, Germany
2. Molecular Biotechnology and Systems Biology, Rheinland-Pfälzische TU Kaiserslautern, Paul-Ehrlich Straße 23, Kaiserslautern D-67663, Germany
3. Leibniz Institute for Plant Genetics and Crop Plant Research (IPK), Corrensstraße 3, Seeland, OT Gatersleben D-06466, Germany