1. Centre for Agri-Environmental Research, School of Agriculture, Policy and Development, University of Reading, Reading RG6 6AR, UK
2. Institute of Integrative and Comparative Biology, University of Leeds, Leeds LS2 9JT, UK
3. Naturalis Biodiversity Centre, 2333 CR Leiden, The Netherlands
4. Centre for Ecology and Hydrology, Lancaster LA1 4AP, UK