1. Department of Zoology, Oregon State University, Cordley Hall 3029, Corvallis, OR 97330, USA
2. School of Biological Sciences, University of Sydney, Heydon-Laurence Building AO8, Science Road, New South Wales 2006, Australia
3. Department of Biology, Texas A&M University, 3258 TAMU, College Station, TX 77843, USA
4. Biology Department, University of Massachusetts, 221 Morrill Science Center, Amherst, MA 01003, USA