1. School of Biological Sciences, The University of Adelaide, Adelaide, SA 5005, Australia
2. Department of Biological Sciences, Mount Holyoke College, South Hadley, MA 01075, USA
3. School of Agriculture, Biomedicine and Environment, La Trobe University, VIC 3086, Australia
4. Museum of Zoology, University of Michigan, Ann Arbor, MI 48108, USA
5. Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI 48109, USA