1. Centre for Biological Diversity, University of St Andrews, St Andrews, Fife KY16 8LB, UK
2. School of Biology, University of St Andrews, St Andrews, Fife KY16 8LB, UK
3. CESAM, Departamento de Biologia, Universidade de Aveiro, Campus de Santiago, 3810 Aveiro, Portugal
4. Department of Life Sciences, The University of the West Indies, St Augustine, Trinidad and Tobago