1. Natural History Museum, University of Oslo, PO Box 1172, Blindern, 0318 Oslo, Norway
2. Integrative Research Center, Field Museum, 1400 South Lake Shore Drive, Chicago IL, 60605, USA
3. Centre for Ecological and Evolutionary Synthesis, Department of Biosciences, University of Oslo, PO Box 1066, Blindern, 0316 Oslo, Norway