1. Max-Planck-Institute for Evolutionary Biology, Research Group for Community Dynamics, August-Thienemann-Straße 2, 24306 Plön, Germany
2. Smithsonian Tropical Research Institute, Apartado Postal 0843-03092, Panamá, República de Panamá
3. GEOMAR Helmholtz Centre for Ocean Research Kiel, Evolutionary Ecology of Marine Fishes, Düsternbrooker Weg 20, 24105 Kiel, Germany
4. Faculty of Mathematics and Natural Sciences, University of Kiel, Christian-Albrechts-Platz 4, 24118 Kiel, Germany