1. DEI, Politecnico di Milano, Via Ponzio 34/5, 20133 Milan, Italy
2. Laboratoire Ecologie et Evolution, CNRS UMR 7625, Ecole Normale Superieure, 46 rue d'Ulm, 75005 Paris, France
3. Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, AZ 85721, USA
4. Evolution and Ecology Program, International Institute for Applied Systems Analysis, 2361 Laxenburg, Austria