1. School of Mathematics and Statistics, University of Sheffield, Sheffield S3 7RH, UK
2. Department of Mathematical Sciences, University of Bath, Claverton Down, Bath BA2 7AY, UK
3. Department of Integrative Biology, University of California Berkeley, Berkeley, CA, USA
4. Department of Mathematics and the Maxwell Institute for Mathematical Sciences, Heriot-Watt University, Edinburgh EH14 4AS, UK
5. Department of Mathematical Sciences, Division of Applied Mathematics, The University of Liverpool, Mathematical Sciences Building, Liverpool L69 7ZL, UK
6. Biosciences, College of Life and Environmental Sciences, University of Exeter, Cornwall Campus, Treliever Road, Penryn, Cornwall TR10 9EZ, UK