1. Biosciences, College of Life and Environmental Sciences; University of Exeter; Cornwall Campus Penryn Cornwall TR10 9EZ United Kingdom
2. Department of Mathematics and the Maxwell Institute for Mathematical Sciences; Heriot-Watt University; Edinburgh EH14 4AS Scotland United Kingdom
3. School of Mathematics and Statistics; University of Sheffield; Sheffield S3 7RH United Kingdom
4. Division of Applied Mathematics; Department of Mathematical Sciences; Mathematical Sciences Building, The University of Liverpool L69 7ZL United Kingdom