1. Conservation Ecology Group, Groningen Institute for Evolutionary Life Sciences, University of Groningen, PO Box 11103, 9700, CC, Groningen, The Netherlands
2. Division of Biological Sciences, University of Montana, 32 Campus Drive, Missoula, MT 59812, USA
3. NIOZ Royal Netherlands Institute for Sea Research, Department of Coastal Systems, Utrecht University, PO Box 59, 1790, AB Den Burg, Texel, The Netherlands
4. Computational Geo-Ecology, Institute for Biodiversity and Ecosystem Dynamics, University of Amsterdam, Sciencepark 904, 1098, XH Amsterdam, The Netherlands