1. Department of Biology, University of Washington, PO Box 351800, Seattle, WA 98195, USA
2. Laboratory of Aquatic Ecology and Evolutionary Biology, Katholieke Universiteit Leuven, Belgium
3. Department of Biological Sciences, University of Notre Dame, Notre Dame, IN, USA
4. Max Planck Institute for Evolutionary Biology, Plön, Germany
5. Department of Ecology and Evolutionary Biology, Cornell University, Ithaca, NY, USA