1. Department of Biology, University of York, Wentworth Way, York YO10 5DD, UK
2. School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich NR4 7TJ, UK
3. Department of Zoology, University of Oxford, Oxford OX1 3PS, UK
4. Department of Genetics, Evolution and Environment, University College London, London WC1E 6BT, UK
5. Institute of Integrative Biology, University of Liverpool, Liverpool L69 7ZB, UK