1. Department of Biological Sciences, Ohio University, Athens, OH 45701, USA
2. Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Invalidenstraße 43 Berlin D-10115, Germany
3. Paleontological Institute and Museum, University of Zurich, Zurich CH-8006, Switzerland
4. Department of Biosciences, Swansea University, Wallace Building, Singleton Park, Swansea SA2 8PP, UK
5. Smithsonian Tropical Research Institute, PO Box 2072, Balboa, Panama