1. Centre for Research into Ecological and Environmental Modelling, University of St Andrews, St Andrews, UK
2. Wildlife Conservation Research Unit, Department of Biology, University of Oxford, Oxford, UK
3. Department of Natural Resources and the Environment, Cornell University, Ithaca, NY, USA
4. Northeast Fisheries Science Center, NOAA National Marine Fisheries Service, Woods Hole, MA, USA
5. Department of Biology, University of Oxford, Oxford, UK
6. Zambian Carnivore Programme, Mfuwe, Zambia