1. Department of Life Sciences, Aberystwyth University, Aberystwyth SY23 3FG, UK
2. Department of Computer Science, Aberystwyth University, Aberystwyth SY23 3DB, UK
3. Centre for Integrative Ecology, School of Life and Environmental Sciences, Deakin University, Waurn Ponds, Victoria 3216, Australia
4. Department of Evolutionary Ecology, National Museum of Natural Sciences, CSIC, 28006 Madrid, Spain
5. International Centre of Insect Physiology and Ecology, PO Box 30772-00100, Nairobi, Kenya