1. Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
2. Institute of Zoology, Zoological Society of London, Regent's Park, London NW1 4RY, UK
3. Hihi Conservation Charitable Trust, Rotorua, New Zealand
4. Helsinki Institute of Life Science (HiLIFE), University of Helsinki, Helsinki 00014, Finland
5. Research program in Organismal and Evolutionary Biology, Faculty of Biological and Environmental Sciences, University of Helsinki, Helsinki 00014, Finland