1. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA 98195, USA
2. Department of Integrative Biology, University of South Florida, Tampa, FL 33620, USA
3. Department of Biology, Depauw University, Greencastle, IN 46135, USA
4. Zoology Department, University of British Columbia, Vancouver, British Columbia, Canada V6K 1H8
5. Hakai Institute, End of Kwakshua Channel, Calvert Island, British Columbia, Canada
6. Department of Ecology, Evolution and Marine Biology, and Marine Science Institute, University of California, Santa Barbara, CA 93106, USA