1. Ecology, Evolution, Environment & Society Graduate Program, Dartmouth College, Hanover, NH 03755, USA
2. Department of Biological Sciences, Dartmouth College, Hanover, NH 03755, USA
3. Department of Mammalogy, Division of Vertebrate Zoology, American Museum of Natural History, New York, NY 10024, USA
4. Smithsonian Tropical Research Institute, Balboa, Ancón, Apartado 0843-03092, Republic of Panama
5. Department of Ecology, Evolution, and Organismal Biology, Brown University, Providence, RI 012912, USA
6. School of Engineering, Brown University, Providence, RI 012912, USA