1. Department of Ecology and Evolutionary Biology, 321 Steinhaus Hall, University of California, Irvine, CA 92697, USA
2. Institute for Advanced Study, Einstein Drive, Princeton, NJ 08540, USA
3. Division of Public Health, Fred Hutchison Cancer Research Center, Seattle, WA 98109, USA
4. Biomathematics Graduate Program, Department of Mathematics, North Carolina State University, Raleigh, NC 27695, USA