1. University Marine Biological Station, Millport, Isle of Cumbrae KA28 0EG, UK
2. IUEM, Ressources halieutiques–Poissons marins, Pl. N. Copernic, Technopôle Brest–Iroise, 29280 Plouzané, France
3. Institute of Marine Research, PO Box 1870 Nordnes, N–5817 Bergen, Norway