1. School of Biological Sciences, University of LiverpoolCrown Street, Liverpool L69 7ZB, UK
2. Facultad de Ciencias Veterinarias, Universidad Nacional del LitoralRP Kreder 2805 3080 Esperanza, Santa Fe, Argentina
3. Global Health Programs, Wildlife Conservation SocietyBronx, NY 10460, USA
4. National Centre for Zoonosis Research, University of LiverpoolChester High Road, Neston CH64 7TE, UK