1. Department of Biological Sciences and NERC Centre for Population Biology, Imperial College London, Silwood Park Campus, Ascot SL5 7PY, UK
2. Molecular Systematics Section, Jodrell Laboratory, Royal Botanic Gardens, Kew TW9 3DS, UK
3. GIS unit, Herbarium, Royal Botanic Gardens, Kew TW9 3AB, UK