1. Department of the Geophysical Sciences, University of ChicagoChicago, IL 60637, USA
2. Department of Plant Sciences, University of TasmaniaHobart, Tasmania 7001, Australia
3. Department of Ecology and Evolutionary Biology, University of TennesseeKnoxville, TN 37996, USA
4. Arnold Arboretum of Harvard UniversityJamaica Plain, MA 02130, USA