1. Department of Biology (area 2), University of YorkPO Box 373, York YO10 5YW, UK
2. Institute of Vertebrate Biology, Academy of Science of the Czech RepublicStudenec 122, 675 02 Konešín, Czech Republic
3. Quercus, School of Biological Sciences, Queen's University Belfast97 Lisburn Road, Belfast BT9 7BL, UK