1. Department of Ecology and Evolutionary Biology, and Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA
2. L. H. Bailey Hortorium, Department of Plant Biology, Cornell University, Ithaca, NY 14853, USA
3. Department of Biology, University of Florida, Gainesville, FL 17 32611, USA
4. Florida Museum of Natural History, University of Florida, Gainesville, FL 17 32611, USA
5. Department of Ecology, Evolution, and Organismal Biology, Iowa State University, Ames, IA 50011, USA