1. Jepson Herbarium and Department of Integrative Biology, 1001 Valley Life Sciences Building #2465, University of California, Berkeley, California 94720‐2465 USA
2. Department of Biological Sciences, University of Pittsburgh, 4249 Fifth Avenue, Pittsburgh, Pennsylvania 15260 USA
3. School of Biological Sciences, King Henry Building, King Henry I Street, University of Portsmouth, Portsmouth PO1 2DY, UK
4. Department of Biology, NTNU, Realfagbygget, Gløshaugen, N‐7491, Trondheim, Norway
5. Institute of Arctic Biology, University of Alaska, P.O. Box 757000, Fairbanks, Alaska 99775‐7000 USA