1. Department of Biology, University of Washington, Seattle, WA, USA
2. Department of Zoology, The University of Melbourne, Victoria 3010, Australia
3. School of Marine and Tropical Biology, James Cook University, Cairns, Queensland 4870, Australia
4. School of Marine and Tropical Biology, James Cook University, Townsville, Queensland 4811, Australia
5. Reef and Rainforest Research Centre, 51 The Esplanade, Cairns, Queensland 4870, Australia