1. Southern Appalachian Biodiversity InstituteRoan Mountain, TN 37687, USA
2. Monte L. Bean Life Science Museum, Brigham Young UniversityProvo, UT 84602, USA
3. Department of Biology, College of CharlestonCharleston, SC 29424, USA
4. Center for Biodiversity and Conservation, Drexel University3141 Chestnut St., Philadelphia, PA 19104, USA
5. Department of Biology, 675 WidtsoeBrigham Young University, Provo, UT 84602, USA