1. Hefei National Laboratory for Physical Sciences at the Microscale, School of Life Sciences, University of Science and Technology of China, Hefei 230027, People's Republic of China
2. Center for Excellence in Brain Science and Intelligence Technology, Chinese Academy of Sciences, Shanghai, 200031, People's Republic of China
3. College of Life Science and Technology, Huazhong University of Science and Technology, Wuhan 430074, People's Republic of China
4. The Lunenfeld-Tanenbaum Research Institute, Mount Sinai Hospital; Department of Molecular Genetics, Department of Physiology, University of Toronto, Toronto, Ontario M5G 1XS, Canada