1. Department of Plant Systems Biology, VIB, Technologiepark 927, 9052 Ghent, Belgium
2. Department of Plant Biotechnology and Genetics, Ghent University, Technologiepark 927, 9052 Ghent, Belgium
3. Division of Plant and Crop Sciences, School of Biosciences, University of Nottingham, Loughborough LE12 5RD, UK