1. Department of Wildland Resources, Utah State University, 5230 Old Main Hill, Logan, UT 84322, USA
2. U.S. Geological Survey, Northern Rocky Mountain Science Center, 2327 University Way, Suite 2, Bozeman, MT 59715, USA
3. U.S. Fish and Wildlife Service, National Elk Refuge, PO Box 510, Jackson, WY 83001, USA
4. Wyoming Game and Fish Department, 432 Mill Street, Pinedale, WY 82941, USA