1. Boyd Orr Centre for Population and Ecosystem Health, Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow G12 8QQ, UK
2. School of Veterinary Medicine, University College Dublin, Dublin, Ireland
3. Royal (Dick) School of Veterinary Studies and Roslin Institute, University of Edinburgh, Easter Bush Campus, Midlothian EH25 9RG, UK
4. Department of Veterinary Services, Ministry of Livestock and Fisheries, Nelson Mandela Road, Dar Es Salaam, Tanzania
5. Department of Infectious Diseases, University of Georgia, Athens, GA 30602, USA
6. Institute of Bioinformatics, University of Georgia, Athens, GA 30602, USA