1. Department of Zoology, University of Cambridge, Cambridge CB2 3EJ, UK
2. CNRS UMR 7205, Muséum National d'Histoire Naturelle, Département Systématique et Evolution, 45 rue Buffon, 75005 Paris, France
3. School of Biosciences, University of Exeter in Cornwall, Penryn TR10 9EZ, UK
4. Institute of Evolutionary Biology, University of Edinburgh, Edinburgh EH9 3JT, UK
5. The Galton Laboratory, University College London, Stephenson Way, London NW1 2HE, UK
6. Wellcome Trust Sanger Institute, Hinxton, Cambridge CB10 1SA, UK
7. The GenePool Genomics Facility, School of Biological Sciences, University of Edinburgh, Edinburgh EH9 3JT, UK