1. Genomics and Computational Biology Graduate Program, University of Pennsylvania, Philadelphia, PA 19103, USA
2. Evolution and Cancer Laboratory, Barts Cancer Institute, Barts and the London School of Medicine and Dentistry, Queen Mary University of London, London EC1M 6BQ, UK
3. Department of Pathology and Laboratory Medicine, University of Pennsylvania, Philadelphia, PA 19103, USA
4. Biodesign Institute, School of Life Sciences, Arizona State University, Tempe, AZ 85287, USA
5. Center for Evolution and Cancer, University of California San Francisco, San Francisco, CA 94143, USA