1. Haerdawei 62, 8854 AC Oosterbierum, The Netherlands
2. Animal Ecology Group, Centre for Ecological and Evolutionary Studies, University of GroningenPO Box 14, 9750 AA Haren, The Netherlands
3. Department of Marine Ecology and Evolution, Royal Netherlands Institute for Sea Research (NIOZ)PO Box 59, 1790 AB Den Burg, Texel, The Netherlands