1. Department of Biology, University of Nevada, Reno, NV 89557, USA
2. Program in Ecology, Evolution and Conservation Biology, University of Nevada, Reno, NV 89557, USA
3. Department of Agriculture, Veterinary and Rangeland Sciences, University of Nevada, Reno, NV 89557, USA
4. United States Fish and Wildlife Service, Klamath Falls Fish and Wildlife Office, Klamath Falls, OR 97602, USA