1. Department of Pharmacology, Perelman School of Medicine, University of Pennsylvania, 36th and Hamilton Walk, Philadelphia, PA 19104, USA
2. Department of Biology, School of Arts and Sciences, University of Pennsylvania, 301 Lynch Laboratories/6018, 433 S. University Avenue, Philadelphia, PA 19104, USA
3. PENN Genome Frontiers Institute, University of Pennsylvania, 301 Lynch Laboratories/6018, 433 S. University Avenue, Philadelphia, PA 19104, USA