1. Centre for Systems Biology at Edinburgh, University of Edinburgh, C. H. Waddington Building, King's Buildings Campus, Mayfield Road, Edinburgh EH9 3JD, UK
2. School of Biological Sciences, University of Edinburgh, King's Buildings Campus, Mayfield Road, Edinburgh EH9 3JH, UK
3. School of Informatics, University of Edinburgh, Informatics Forum, 10 Crichton Street, Edinburgh EH8 9AB, UK