1. Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, PA 19085, USA
2. Thackeray Wildlife Foundation, Vaibhav Chambers, Bandra, Mumbai 400051, India
3. Museu de História Natural e da Ciência da Universidade do Porto, Praça Gomes Teixeira, 4099-002 Porto, Portugal