1. Department of Genetics, Evolution and Environment, Centre for Biodiversity and Environment Research, Darwin Building, UCL, Gower Street, London WC1E 6BT, UK
2. School of Biological Sciences and the Environment Institute, The University of Adelaide, Adelaide, South Australia 5005, Australia
3. Institute of Zoology, Zoological Society of London, Regent's Park, London NW1 4RY, UK
4. Distinguished Scientist Fellowship Program, King Saud University, Riyadh 1145, Saudi Arabia